We carried out a hive inspection over the weekend and as predicted, the problem hive had died. It was full of dead bees clustered together. We had a close look at the bees and they did not appear deformed. That should mean it wasn't an infection or varroa parasites. It may be that it starved to death. This sometimes happens even though there is food available elsewhere in the hive.
The dead hive has been cleared away and will be used for the next swarm. All other hives on the apiary were checked and were in good order. We added supers to 2.
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